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There are 1391 sixteen-letter words containing G, L, N, R and S

(page 9)

tiger␣salamanderstools␣of␣ignoranceToronto␣blessingstracheosyringealtraditionalisingtraffic␣signal␣boxtranquillisinglytranscategoricaltransclassifyingtransdiaphragmaltransgeneticallytransglucosidasetransglucosylasetransglucosylatetransglutaminasetransglycosidasetransglycosylasetransglycosylatetranslanguagingstranslatologicaltranslatologiststransligamentoustranslingualismstranslogarithmictransmetallatingtransmigrationaltransoesophagealTranspadane␣Gaulstransphonologisetransphonologizetranssexualizingtransverbalizingtravelling␣cranestread␣on␣eggshellstrembling␣poplarstriangular␣colonstriangular␣prismstrichalcogenidestrichostrongylestrichostrongylidtrigeminal␣nervestrigonocephaliestriple␣energizersTurks␣Gut␣Long␣Pondturning␣the␣tablestwilight␣industryultracentrifugesultrahomogeneousultrasimplifyingultrasonographerultrasonographicultrasonologicalultrasonologistsultrasonosurgeryuncompromisinglyuncopyrightablesunderclusteringsundergeneralisedundergeneralisesundergeneralizesunderstimulatingundertranslatingundiscourageableunenterprisinglyunforgivablenessungovernablenessuniversal␣algebrauniversal␣bankinguniversal␣grammaruniversal␣grinderunneighborlinessunreflectingnessunregistrabilityunrevengefulnessunscrutinizinglyunspiritualisingunspiritualizingunstraightenableuntransfigurableupward␣lightningsurban-rural␣gminasurogynaecologisturogynecologistsUS␣Virgin␣Islanderutilitarianisingvaginal␣dischargevaginal␣processesvaingloriousnessvan␣der␣Waals␣gasesvariable␣bindingsvegetable␣gardensvegetable␣rennetsvertical␣kampongsvertical␣kampungsvictualling␣yardsVirginia␣wild␣ryesvirginity␣pledgesvirtue␣signallersvirtue-signalling virtue␣signallingvisiting␣lecturervisiting␣scholarsvomeronasal␣organwalking␣marriageswalking␣simulatorwalking␣the␣boardswandering␣spleenswarning␣trianglesWartenberg␣wheelsweathering␣steelsWestern␣astrologyWeststellingwerfwheels␣are␣turningwhiptail␣stingraywhistling␣marmotswholegrain␣breads whole-grain␣breadsWigner-Seitz␣cellswildlife␣crossingWillmore␣energieswillow␣ptarmigansWinchester␣gallonwishful␣recyclingwith␣a␣grain␣of␣saltwith␣flying␣colorsWivelsfield␣Greenwoolly␣fringe␣mossworking-classizedworking-classizesworking␣languagesxanthogranulomasyellow␣supergiantzero-shot␣learning

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